Andrea "Andy" Sachs, a recent Brown graduate hired as junior personal assistant to a powerful and tyrannical fashion magazine editor., Miranda Priestly, British-born (as Miriam Princhek) editor-in-chief of Runway, an influential fashion magazine published by the Elias-Clark company. She is known for wearing a white Hermès scarf somewhere on her person every day, and treats her subordinates in a manner that borders on emotional and psychological abuse, Emily Charlton, Miranda's former junior assistant, now her senior assistant. She and Andrea have a conflicted relationship., Alex Fineman, Andrea's boyfriend, who teaches at an elementary school in the South Bronx through Teach for America., Lily Goodwin, a free-spirited graduate student in Russian literature at Columbia with curly black hair. She is Andrea's roommate., Hunter Tomlinson, a prominent New York tax attorney who is Miranda's current husband, Caroline and Cassidy, the twin daughters Miranda dotes on., Benjamin, He is Lily's ex-boyfriend. He was involved in the car accident with Lily.

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